Cheumatopsyche

?
*Cheumatopsyche*
? Cheumatopsyche
	Cheumatopsyche columnata
Научная классификация
Царство:	Животные
Тип:	Членистоногие
Класс:	Насекомые
Отряд:	Ручейники
Семейство:	Гидропсихиды
Род:	Cheumatopsyche
Латинское название
	Cheumatopsyche Wallengren, 1891
Синонимы
	Hydropsychodes Ulmer, 1905; Ulmeria Navás, 1918; Ecnopsyche Banks, 1913; Plectropsyche Ross, 1947;
ITIS	115408
NCBI	177865

Cheumatopsyche (лат.) — род ручейников семейства гидропсихиды (Hydropsychidae) подотряда Annulipalpia.

Распространение

Всесветное^[1].

Описание

Мелкие ручейники, длина тела менее 1 см. В основном коричневого или жёлтого цвета, а некоторые виды — тёмные с характерным белым рисунком. Помимо характерной комбинации основных родовых признаков Cheumatopsyche легко отличить от всех других родов гидропсихин по характерной генитальной структуре. Только, возможно, предковая группа видов Cheumatopsyche pali, населяющая Мадагаскар и окрестности имеет удлинённые преанальные придатки. У всех остальных групп видов преанальные придатки меньше. Другими явно примитивными состояниями первичных родовых признаков в этой группе являются: наличие в задних крыльях m-cu; формула шпор 244; и наличие вилок 1235 задних крыльев. Существует 3 производных признака: проэпистернальная щетинистая бородавка отсутствует; близко расположенные поперечные жилки передних крыльев от m-cu до cu; открытая медиальная ячейка заднего крыла. На голове 5-9 щетинок бородавок. Претарзальные коготки самцов имеют разную форму, асимметричны, у некоторых видов с боков обрамлены пучком щетинок. Однако есть виды, в основном с Мадагаскара, с идеально симметричными претарзальными когтями на всех ногах. Формула шпор 2-4-4^[1].

Личинка

Систематика

Около 350 видов. Род был впервые выделен в 1891 году на основании типового вида Hydropsyche lepida Pictet, 1834. Cheumatopsyche близок к Potamyia, но жилки задних крыльев M и Cu1 не параллельны и не близки друг к другу у Cheumatopsyche, а в гениталиях язычковая мезоапикальная доля хорошо развита у нескольких групп видов Cheumatopsyche и никогда не присутствует у Potamyia. Валидный статус рода подтверждён в ходе ревизии, проведённой в 2008 году венгерским энтомологом János Oláh (Department of Environmental Management, Tessedik College, Сарваш, Венгрия) с коллегами^[1]^[2]^[3]^[4]^[5].

C. abhugna (J Olah & KA Johanson, 2008)
C. acuminata (J Olah & KA Johanson, 2008)
C. addita (L Navas, 1916)
C. admetos (H Malicky & P Chantaramongkol, 1996)
C. agha (J Olah & KA Johanson, 2008)
C. agnetae (J Olah & KA Johanson, 2008)
C. akana (B Statzner, 1984)
C. akisena (J Olah & KA Johanson, 2008)
C. alampeta (A Neboiss, 2002)
C. albocincta (Banks, 1937)
C. albofasciata (R McLachlan, 1872)
C. albomaculata (Ulmer, 1905)
C. alfierii (L Navas, 1927)
C. alvarezi (W Wichard, MM Solorzano-Kraemer, C Luer, 2006)
C. ambala (J Olah & KA Johanson, 2008)
C. ambodivoa (J Olah & KA Johanson, 2008)
C. ambohima (J Olah & KA Johanson, 2008)
C. amboinica (L Navas, 1928)
C. amdakaramaya (J Olah & KA Johanson, 2008)
C. ampanga (J Olah & KA Johanson, 2008)
C. amurensis (AV Martynov, 1934)
C. analis (Banks, 1903)
C. andohahela (J Olah & KA Johanson, 2008)
C. anema (G Marlier, 1959)
C. angusta (Ulmer, 1930)
C. anonima (J Olah & KA Johanson, 2008)
C. anthracias (W Mey, 1996)
C. antoniensis (H Malicky, 1982)
C. aphanta (HH Ross, 1938)
C. apicata (L Navas, 1931)
C. arizonensis (S Ling, 1938)
C. asaha (J Olah & KA Johanson, 2008)
C. aterrima (G Marlier, 1961)
C. automedon (H Malicky & P Chantaramongkol, 1997)
C. ayopa (J Olah & KA Johanson, 2008)
C. balaromta (J Olah & KA Johanson, 2008)
C. banaueensis (W Mey, 1995)
C. banksi (Mosely, 1942)
C. barakambra (J Olah & KA Johanson, 2008)
C. bardiana (H Malicky, 1997)
C. bhatrapura (H Malicky, 1979)
C. bibbensis (Gordon, Harris & Lago, 1986)
C. bidikala (J Olah & KA Johanson, 2008)
C. bifida (B Statzner, 1975)
C. bimaculata (Ulmer, 1930)
C. boettgeri (B Statzner, 1975)
C. bohakayi (J Olah & KA Johanson, 2008)
C. bolosa (J Olah & KA Johanson, 2008)
C. brazzana (J Olah & KA Johanon, 2008)
C. brevilineata (M Iwata, 1927)
C. brevis (Ulmer, 1930)
C. brunnea (S Jacquemart, 1961)
C. burgeonia (L Navas, 1931)
C. burksi (HH Ross, 1941)
C. cahaba (Gordon, Harris & Lago, 1986)
C. caieta (H Malicky & P Chantaramongkol, 1997)
C. calawagana (W Mey, 1995)
C. camena (H Malicky, 1997)
C. camerunica (J Olah & KA Johanson, 2008)
C. camilla (H Malicky, 1997)
C. campyla (HH Ross, 1938)
C. capitata (L Navas, 1936)
C. capitella (AV Martynov, 1927)
C. caprotina (H Malicky, 1997)
C. carmentis (H Malicky & P Chantaramongkol, 1997)
C. carna (H Malicky & P Chantaramongkol, 1997)
C. ceramensis (W Mey, 1999)
C. ceres (H Malicky & P Chantaramongkol, 1997)
C. clavalis (AV Martynov, 1930)
C. cocles (H Malicky & P Chantaramongkol, 1997)
C. cognita (Ulmer, 1951)
C. columnata (AV Martynov, 1935)
C. comis (SW Arnold & CR Arnold, 1961)
C. comorina (L Navas, 1931)
C. concava (Ulmer, 1930)
C. concordia (H Malicky, 1997)
C. congolana (J Olah & KA Johanson, 2008)
C. contexta (Ulmer, 1951)
C. copia (H Malicky & P Chantaramongkol, 1997)
C. copiosa (DE Kimmins, 1956)
C. cornix (H Malicky, 1997)
C. costalis (Banks, 1913)
C. cressida (H Malicky, 1997)
C. criseyde (H Malicky & P Chantaramongkol, 1997)
C. curvata (AV Martynov, 1935)
C. chariklo (H Malicky & P Chantaramongkol, 1997)
C. charites (H Malicky & P Chantaramongkol, 1997)
C. charybdis (H Malicky, 1997)
C. cheesmanae (DE Kimmins, 1962)
C. chihonana (M Kobayashi, 1987)
C. chimaira (H Malicky, 1997)
C. chinensis (AV Martynov, 1930)
C. chione (H Malicky & P Chantaramongkol, 1997)
C. chirapali (J Olah & KA Johanson, 2008)
C. chitawana (J Olah & KA Johanson, 2008)
C. chlorogastra (L Navas, 1932)
C. chryseis (H Malicky & P Chantaramongkol, 1997)
C. chrysothemis (H Malicky & P Chantaramongkol, 1997)
C. danae (H Malicky, 1998)
C. davisi (J Olah & KA Johanson, 2008)
C. deani (A Neboiss, 2002)
C. delamarei (S Jacquemart, 1965)
C. dhanikari (H Malicky, 1979)
C. diehli (H Malicky, 1997)
C. digitata (Mosely, 1935)
C. diminuta (Walker, 1852)
C. doan (J Olah & KA Johanson, 2008)
C. dostinei (JC Dean, 2001)
C. dubitans (Mosely, 1942)
C. duma (H Malicky, 2009)
C. ebal (H Malicky, 2009)
C. ecsedii (J Olah & KA Johanson, 2008)
C. edista (AE Gordon, 1974)
C. edom (H Malicky, 2009)
C. ehi (H Malicky, 2009)
C. ekona (J Olah & KA Johanson, 2008)
C. ela (DG Denning, 1942)
C. emas (CJ Geraci, 2008)
C. engor (J Olah & PC Barnard, 2008)
C. enigma (Morse, Ross & Gordon, 1971)
C. enonis (HH Ross, 1938)
C. erawan (H Malicky & P Chantaramongkol, 2009)
C. ernestheissi (H Malicky, 1997)
C. etrona (HH Ross, 1941)
C. excisa (Ulmer, 1930)
C. expeditionis (Ulmer, 1938)
C. explicanda (B Statzner, 1975)
C. fahara (J Olah & KA Johanson, 2008)
C. falcifera (Ulmer, 1930)
C. fansipangensis (W Mey, 1996)
C. fausta (L Navas, 1936)
C. fianara (J Olah & KA Johanson, 2008)
C. flavellata (W Mey, 2004)
C. flavosulphurea (W Mey, 1998)
C. fusca (L Navas, 1936)
C. gaia (H Malicky, 1997)
C. galahittigama (F Schmid, 1958)
C. galapitikanda (F Schmid, 1958)
C. galumata (J Olah & KA Johanson, 2008)
C. galloisi (S Matsumura, 1931)
C. garulupota (Olah et al., 2007)
C. gelita (DG Denning, 1952)
C. geora (DG Denning, 1948)
C. georgulmeri (W Mey, 1998)
C. gibbsi (B Statzner, 1984)
C. globosa (Ulmer, 1910)
C. gordonae (PK Lago & SC Harris, 1983)
C. gracilis (Banks, 1899)
C. guadunica (Y Li, 1988)
C. guerneana (L Navas, 1916)
C. gyra (HH Ross, 1938)
C. hageni (Olah et al., 2007)
C. halima (DG Denning, 1948)
C. harwoodi (DG Denning, 1948)
C. hasalaka (J Olah & KA Johanson, 2008)
C. helma (HH Ross, 1939)
C. hippolyte (H Malicky, 1997)
C. hoasena (J Olah & KA Johanson, 2008)
C. holzschuhi (H Malicky, 1997)
C. hoogstraali (HH Ross, 1947)
C. imatso (J Olah & KA Johanson, 2008)
C. incomptaria (W Mey, 1998)
C. indica (L Navas, 1932)
C. infascia (AV Martynov, 1934)
C. jacquemarti (J Olah & KA Johanson, 2008)
C. jaekam (H Malicky & P Chantaramongkol, 2009)
C. japonica (L Navas, 1916)
C. jiriana (H Malicky, 1997)
C. joariva (J Olah & KA Johanson, 2008)
C. junolahi (J Olah & KA Johanson, 2008)
C. kahat (H Malicky & P Chantaramongkol, 2009)
C. kakaduensis (JC Dean, 2001)
C. kala (J Olah & KA Johanson, 2008)
C. kali (CJ Geraci, 2008)
C. kebumena (H Malicky, 1997)
C. kim (J Olah & KA Johanson, 2008)
C. kimminsi (G Marlier, 1961)
C. kindamba (J Olah & KA Johanson, 2008)
C. kinlockensis (Gordon et al., 1986)
C. kirimaduwa (F Schmid, 1958)
C. kissi (G Marlier, 1961)
C. kraepelini (Ulmer, 1905)
C. krugerana (J Olah & KA Johanson, 2008)
C. kumasa (J Olah & KA Johanson, 2008)
C. kuranishii (J Olah & KA Johanson, 2008)
C. kysonia (J Olah & KA Johanson, 2008)
C. langsonina (L Navas, 1933)
C. lasia (HH Ross, 1938)
C. lateralis (KH Barnard, 1934)
C. lebasi (L Navas, 1932)
C. leleupi (G Marlier, 1961)
C. leloupi (S Jacquemart, 1957)
C. lepida (F.J. Pictet, 1834)
C. lesnei (Mosely, 1932)
C. lestoni (DG Gibbs, 1973)
C. lobata (G Marlier, 1943)
C. logani (AE Gordon & SD Smith, 1974)
C. lolo (J Olah & KA Johanson, 2008)
C. longiclasper (Y Li, 1988)
C. longinosnavasi (W Mey, 1998)
C. lucida (Ulmer, 1907)
C. macentae (Gibon, Guenda & Coulibaly, 1994)
C. maculata (Mosely, 1934)
C. maculipennis (Ulmer, 1930)
C. madagassa (L Navas, 1923)
C. mafana (J Olah & KA Johanson, 2008)
C. mahakaya (J Olah & KA Johanson, 2008)
C. malmi (J Olah & KA Johanson, 2008)
C. mandrara (J Olah & KA Johanson, 2008)
C. mariannae (J Olah & KA Johanson, 2008)
C. marieni (S Jacquemart, 1965)
C. marmorata (L Navas, 1922)
C. masia (L Navas, 1920)
C. masiposa (J Olah & KA Johanson, 2008)
C. mattheei (W Mey, 1992)
C. matuta (H Malicky, 2004)
C. mendolonga (J Olah & KA Johanson, 2008)
C. meo (J Olah & KA Johanson, 2008)
C. meruana (L Navas, 1934)
C. meyi (H Malicky, 1997)
C. mickeli (DG Denning, 1942)
C. minuscula (Banks, 1907)
C. mixta (W Mey, 2003)
C. modica (R McLachlan, 1871)
C. mollala (HH Ross, 1941)
C. montapo (J Olah & KA Johanson, 2008)
C. morsei (AE Gordon, 1974)
C. musiana (Ulmer, 1951)
C. namha (J Olah & KA Johanson, 2008)
C. naphis (H Malicky & P Chantaramongkol, 2009)
C. nathanbanksi (W Mey, 1998)
C. naumanni (H Malicky, 1986)
C. niasensis (H Malicky, 1997)
C. nigrescens (L Navas, 1932)
C. ningmapa (F Schmid, 1975)
C. nohondoan (J Olah & KA Johanson, 2008)
C. nondra (J Olah & KA Johanson, 2008)
C. nubila (DE Kimmins, 1963)
C. obscurata (Ulmer, 1930)
C. obtusa (S Jacquemart, 1963)
C. okinawana (J Olah & KA Johanson, 2008)
C. oktedit (J Olah & KA Johanson, 2008)
C. opposita (Banks, 1931)
C. oxa (HH Ross, 1938)
C. padaha (J Olah & KA Johanson, 2008)
C. pali (J Olah & KA Johanson, 2008)
C. pallida (Banks, 1920)
C. paracostalis (W Mey, 2003)
C. parafra (J Olah & KA Johanson, 2008)
C. parentum (AE Gordon, 1974)
C. pasella (HH Ross, 1941)
C. peirithoos (H Malicky & N Changthong, 2005)
C. persica (W Mey, 2004)
C. petersi (Ross, Morse & Gordon, 1971)
C. pfundsteini (B Statzner, 1984)
C. piljanae (W Mey, 1999)
C. pinaca (HH Ross, 1941)
C. pinula (DG Denning, 1952)
C. pison (H Malicky & P Chantaramongkol, 2009)
C. pitella (DG Denning, 1968)
C. plutonis (Banks, 1913)
C. praepilata (CJ Geraci, 2008)
C. processuata (AV Martynov, 1927)
C. pulchripennis (Banks, 1939)
C. pulverulenta (DG Gibbs, 1973)
C. pulla (L Navas, 1918)
C. punctata (S Jacquemart, 1961)
C. quadrifasciata (J Olah & KA Johanson, 2008)
C. ranoma (J Olah & KA Johanson, 2008)
C. ranomafana (J Olah & KA Johanson, 2008)
C. rantsoa (J Olah & KA Johanson, 2008)
C. reticulata (Banks, 1913)
C. richardsoni (AE Gordon, 1974)
C. rienga (J Olah & KA Johanson, 2008)
C. rineta (J Olah & KA Johanson, 2008)
C. robisoni (SR Moulton & KW Stewart, 1996)
C. robusta (Walker, 1852)
C. roscida (L Navas, 1934)
C. rossi (AE Gordon, 1974)
C. rubachi (W Mey, 1995)
C. sagitta (M Kobayashi, 1987)
C. saiyok (H Malicky & P Chantaramongkol, 2009)
C. saltorum (W Mey, 1998)
C. sambava (J Olah & KA Johanson, 2008)
C. sandrae (J Olah & KA Johanson, 2008)
C. saplaca (J Olah & KA Johanson, 2008)
C. sauteri (L Navas, 1933)
C. schmidiana (Olah et al., 2007)
C. schwendingeri (H Malicky & P Chantaramongkol, 1997)
C. septa (J Olah & KA Johanson, 2008)
C. sessilis (G Marlier, 1961)
C. sexfasciata (Ulmer, 1904)
C. simayorum (J Olah & KA Johanson, 2008)
C. simplex (DE Kimmins, 1963)
C. sipitanga (J Olah & KA Johanson, 2008)
C. smithi (AE Gordon, 1974)
C. socia (L Navas, 1927)
C. songbua (J Olah & KA Johanson, 2008)
C. songda (J Olah & KA Johanson, 2008)
C. sordida (HA Hagen, 1861)
C. speciosa (Banks, 1904)
C. spinifera (S Jacquemart, 1967)
C. spinosa (F Schmid, 1959)
C. stenocyta (L Navas, 1932)
C. stigma (DE Kimmins, 1955)
C. suffusa (L Navas, 1932)
C. surgens (Y Li & L Tian, 1990)
C. suswanad (J Olah & KA Johanson, 2008)
C. suteri (JC Dean, 2001)
C. taipeiana (M Kobayashi, 1987)
C. tanana (J Olah & KA Johanson, 2008)
C. tanidai (J Olah & KA Johanson, 2008)
C. tectifera (G Marlier, 1959)
C. telensis (H Malicky, 1997)
C. temburonga (H Malicky, 1997)
C. tenerrima (G Marlier, 1961)
C. tenga (CJ Geraci, 2008)
C. thaba (Mosely, 1948)
C. theophane (H Malicky & T Prommi, 2006)
C. thomasseti (Ulmer, 1931)
C. timgulile (J Olah & KA Johanson, 2008)
C. tincta (L Navas, 1932)
C. tinjar (DE Kimmins, 1955)
C. tokunagai (M Tsuda, 1940)
C. tournii (Gibon, Guenda & Coulibaly, 1994)
C. tramota (H Malicky & P Chantaramongkol, 1997)
C. transmutata (W Mey, 1998)
C. triangularis (Ulmer, 1931)
C. trifascia (Y Li, 1988)
C. trifida (Mosely, 1935)
C. trilari (H Malicky & P Chantaramongkol, 1997)
C. truncata (AV Martynov, 1935)
C. tungpa (J Olah & KA Johanson, 2008)
C. tunmodera (Olah et al., 2007)
C. uchidai (M Kobayashi, 1987)
C. ulmeri (L Navas, 1933)
C. unicalcarata (W Mey, 1992)
C. usambara (J Olah & KA Johanson, 2008)
C. vacoana (J Olah & KA Johanson, 2008)
C. vala (H Malicky, 1992)
C. vannotei (AE Gordon, 1974)
C. varia (DE Kimmins, 1955)
C. ventricosa (Y Li & D Dudgeon, 1988)
C. villosa (L Navas, 1934)
C. virginica (DG Denning, 1949)
C. wabasha (DG Denning, 1948)
C. wellsae (JC Dean, 2001)
C. wrighti (HH Ross, 1947)
C. yangmorseorum (J Olah & KA Johanson, 2008)
C. zao (J Olah & KA Johanson, 2008)

Примечания

↑ ¹ ² ³ Oláh J., Johanson K. A. Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae) (англ.) // Zootaxa : Журнал. — Auckland, New Zealand: Magnolia Press, 2008. — Vol. 1802. — P. 1—248. — ISBN 978-1-86977-222-2. — ISSN 1175-5326. — doi:10.5281/zenodo.5126123.
↑ Olah, J., Flint, O. S., Jr., Johanson, K. A. 2007. Cheumatopsyche galapitikanda species cluster in Sri Lanka with the description of four new species (Trichoptera: Hydropsychidae). Braueria, 34, 21 — 28.
↑ Ross, H. H. (1947) Descriptions and records of North American Trichoptera, with synoptic notes. Transactions of the American Entomological Society, 73, 125—168.
↑ Pandher, M.S., Malicky, H. & Parey, S.H. 2018. New species of the genus Cheumatopsyche Wallengren 1891 (Insecta: Trichoptera: Hydropsychidae) from Indian Himalaya. Zootaxa 4379(3): 407—420. DOI: 10.11646/zootaxa.4379.3.4
↑ Thawarorit, K.; Sangpradub, N.; Morse, J.C. 2013: Five new species of the genus Cheumatopsyche (Trichoptera: Hydropsychidae) from the Phetchabun Mountains, Thailand. Zootaxa 3613(5): 445—454. DOI: 10.11646/zootaxa.3613.5.2

Ссылки

[Oláh2008-1] ¹ ² ³ Oláh J., Johanson K. A. Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae) (англ.) // Zootaxa : Журнал. — Auckland, New Zealand: Magnolia Press, 2008. — Vol. 1802. — P. 1—248. — ISBN 978-1-86977-222-2. — ISSN 1175-5326. — doi:10.5281/zenodo.5126123.

[2] Olah, J., Flint, O. S., Jr., Johanson, K. A. 2007. Cheumatopsyche galapitikanda species cluster in Sri Lanka with the description of four new species (Trichoptera: Hydropsychidae). Braueria, 34, 21 — 28.

[3] Ross, H. H. (1947) Descriptions and records of North American Trichoptera, with synoptic notes. Transactions of the American Entomological Society, 73, 125—168.

[4] Pandher, M.S., Malicky, H. & Parey, S.H. 2018. New species of the genus Cheumatopsyche Wallengren 1891 (Insecta: Trichoptera: Hydropsychidae) from Indian Himalaya. Zootaxa 4379(3): 407—420. DOI: 10.11646/zootaxa.4379.3.4

[5] Thawarorit, K.; Sangpradub, N.; Morse, J.C. 2013: Five new species of the genus Cheumatopsyche (Trichoptera: Hydropsychidae) from the Phetchabun Mountains, Thailand. Zootaxa 3613(5): 445—454. DOI: 10.11646/zootaxa.3613.5.2

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